0:04
the lower Pline period spanning from
0:07
about 2.6 million to 780,000 years ago
0:12
was a time of remarkable Evolution and
0:17
hominin at the heart of this fascinating
0:19
chapter are some truly intriguing
0:22
figures like Homo erectus and Homo
0:25
antecessor early in this period around
0:28
1.8 million years ago Homo erectus
0:31
emerged and became the dominant hominin
0:34
species in Africa and parts of Asia
0:37
known for their use of more advanced
0:39
tools and their ability to adapt to
0:41
diverse environments Homo erectus marked
0:44
a significant step in human
0:47
evolution homo antecessor represents one
0:50
of the earliest known human species in
0:52
Western Europe homo antecessor showed a
0:55
mix of primitive and more modern traits
0:58
indicating a transitional phase in human
1:02
evolution this species was likely
1:04
involved in early tool making and social
1:06
behaviors that would set the stage for
1:10
advancements homo antecessor offers a
1:13
glimpse into human exploration of the
1:15
vast Landscapes of Europe during the
1:17
lower ply toine Epoch they were crafting
1:21
rudimentary tools navigating shifting
1:24
environments and slowly shaping the
1:28
history but the homo antecessor is one
1:31
of the earliest known species in Europe
1:33
and provides crucial evidence of the
1:35
technological and behavioral
1:37
advancements that marked this
1:39
era with a mix of primitive and more
1:42
advanced traits these early humans were
1:45
not just surviving but beginning to
1:47
demonstrate complex behaviors their
1:49
tools were simple yet Innovative for
1:52
their time reflecting their growing
1:54
cognitive abilities and
1:57
adaptability homo antecessor fossils
1:59
were first discovered at a site called
2:01
Grand Delina located in Sierra deap pora
2:06
Spain the Sierra de atap pora is a small
2:10
mountain range in Northern Spain located
2:15
bergus it covers an area of about 25
2:18
Square km and has a maximum height of
2:24
M the Sierra is situated between two
2:27
river basins the duero and the e
2:30
and is an important natural feature
2:34
regions the area experiences both
2:36
Atlantic and Mediterranean climates
2:39
which have likely contributed to its
2:41
Rich biodiversity making it an
2:43
attractive place for ancient human
2:45
ancestors during the Pline and Holocene
2:49
periods many important archaeological
2:52
and paleontological sites are found in
2:54
the Sierra di atap porca particularly
2:57
within a system of caves formed in the
3:01
rocks these fossils were found between
3:04
1994 and 1996 during archaeological
3:08
excavations the site has many layers
3:10
named td1 to td11 with the fossils
3:14
coming from one specific layer called
3:17
td6 the td6 level has been divided into
3:20
three sublevels named td61 td6 2 and TD
3:25
63 the human fossils together with more
3:29
than 300 artifacts and several thousand
3:31
micro and macro mammal fossil remains
3:34
come from sublevel td6
3:37
2 the fossils were found to be from a
3:40
time between 790,000 and 960,000 years
3:44
ago during the early Pline
3:47
period fossilized pollen suggests a
3:50
Mediterranean climate in td6 and the
3:53
study of amphibians and reptiles
3:55
indicates slightly warmer temperatures
4:01
the human remains found in td6 of the
4:03
grand dolina cave are well preserved but
4:06
many are in fragments due to evidence of
4:09
cannibalism researchers studied various
4:11
features of the skulls and bones
4:14
including teeth Jaws in the
4:17
face in trying to determine the minimum
4:20
number of individuals represented in td6
4:23
researchers face challenges due to the
4:25
extensive fragmentation of The
4:28
Remains the T D6 collection includes
4:31
deciduous teeth permanent inzer germs
4:34
and complete lower incizors suggesting
4:37
the presence of both immature and adult
4:41
individuals the estimation of minimum
4:43
number of individuals is preliminary but
4:46
it's notable that a high percentage of
4:48
about 75% of the individuals are
4:52
immature one important fossil from the
4:55
grand alen site is a parietal bone from
4:58
a young individual estimated to be
5:01
between 4 and 12 years old this bone
5:04
shows some primitive features similar to
5:07
earlier human species like homo orter
5:11
erectus the shape of the bone suggests
5:14
the individual would carry these early
5:18
adulthood in contrast the facial Bones
5:22
from the same site show more modern
5:25
features for instance the midface of one
5:28
specimen resembles modern humans with a
5:30
distinct canine fossa and a specific
5:35
jaw these traits are similar to what we
5:37
see in today's humans but were present
5:40
earlier than in most other known
5:43
fossils the mandibles from grelina are
5:46
notably lighter and less robust compared
5:52
fossils they also have some primitive
5:54
features but show some similarities to
5:56
modern humans and Neanderthals in
5:59
specific as aspects like the shape of
6:01
the mandibular Notch and the absence of
6:06
features the teeth of these early humans
6:09
have a mix of primitive and derived
6:11
traits for example the teeth show
6:13
patterns similar to both early humans
6:16
and Neanderthals but also some unique
6:18
features that differentiate
6:21
them overall these fossils from Grand
6:24
Delina show a mix of old and new traits
6:27
this combination of features helps
6:29
scientists understand how human
6:31
evolution happened showing both
6:33
primitive and modern characteristics in
6:37
ancestors in Europe the earliest
6:39
technological record is represented by
6:41
the scarce and Scattered mode one
6:44
technology often deriving from
6:46
occasional occupations or restricted
6:49
areas the tools and materials found at a
6:52
site do not provide a complete picture
6:55
of the actual activities and
6:57
technological processes that occurred
7:01
the earliest tools have been found at a
7:05
lefante people at this site used mostly
7:08
simple stone Flakes and some Limestone
7:10
pieces to make tools these tools were
7:13
created using a straightforward
7:14
technique where flakes were struck off
7:18
Direction the site was used occasionally
7:21
mainly for making tools and eating
7:23
animal food they might have done these
7:26
activities either inside the cave or
7:30
the tools found at the SEMA Delante site
7:33
are associated with early human species
7:35
but it's not definitively established
7:38
that they were made by homo
7:41
antecessor the site is older than the
7:43
earliest known homo antecessor fossils
7:46
which are found at Grand
7:47
Delina at SEMA Del elephante the tools
7:51
are typically linked to early hominins
7:54
from around 1.2 million years
7:59
especially in the td6 level the tool
8:02
making techniques became more advanced
8:05
people used various materials like chirt
8:07
and quartzite to make tools and employ
8:09
different techniques to shape
8:12
them they created a variety of tools and
8:15
while they used many small and
8:17
medium-sized ones there were no large
8:19
tools for heavy tasks this indicates
8:22
that their tool making and usage were
8:24
becoming more sophisticated and
8:28
specialized the most one technology at
8:30
td6 of grand Alina homo antecessor
8:33
demonstrated adaptable tool making by
8:36
using various raw materials like small
8:38
uniform Cretaceous chir and quartz while
8:41
less carefully selecting quartzite and
8:44
Sandstone they employed different core
8:47
reduction strategies such as early
8:49
discarding or extensive shaping by
8:52
rotating the core depending on the
8:56
quality this flexibility in practi iCal
8:59
approach rather than a rigid plan
9:02
suggests they adapted their techniques
9:04
to the specific materials and conditions
9:08
encountered the site was used much more
9:10
intensively during the lower PTO scene
9:13
suggesting it was a regular home base
9:15
for Gathering large mammals in the
9:18
cave these included complex behaviors
9:22
including hunting carcass transport
9:24
social cooperation and food sharing as
9:27
well as the oldest evidence of cannibal
9:32
prehistory they mainly hunted
9:34
opportunistically as seen from the cut
9:36
marks and breakage patterns on Bones
9:39
which indicate they often had Early
9:41
Access to their prey and transported
9:43
intact carcasses back to their
9:46
base the choices made in transporting
9:49
these carcasses were influenced by
9:51
various factors not just the weight of
9:53
the animals while some bones are missing
9:56
from the fossil record due to carnivore
9:58
activity it's clear that homo antecessor
10:01
was responsible for processing the
10:04
carcasses multiple individuals
10:06
participated in hunting and transporting
10:09
carcasses showing signs of social
10:12
cooperation food sharing and a division
10:17
group to understand the position of homo
10:20
antecessor in human evolution it is
10:23
worthwhile to discuss homo
10:26
heidelbergensis the concept of homo hyle
10:28
bensis start started in 1983 with the
10:31
study of a skull from petralona
10:34
Greece over time scientists have debated
10:37
whether this species originated in
10:39
Africa or Western Asia and whether it
10:41
might be the common ancestor of modern
10:46
Neanderthals the name Homo heidleberg
10:48
insis comes from a mandible found near
10:50
heidleberg Germany which was dated to
10:53
around 616,000 years ago this suggests
10:57
it could be the oldest known European
11:01
human in the period between 770,000 and
11:06
126,000 years ago Earth was buzzing with
11:09
action this period called the middle
11:12
Pline was a pivotal era in human history
11:16
it witnessed the rise of various human
11:18
species across different parts of the
11:21
world modern humans emerged in Africa
11:25
while neanderthals made their Mark in
11:26
Europe and the denisovans left their
11:31
Asia alongside Advanced species more
11:34
primitive features were seen in Homo
11:38
homoni and earlier species like homo
11:41
floresiensis in Flores island of
11:43
Indonesia and Homo lanis in the
11:46
Philippines added to the
11:49
diversity during the middle PTO scene
11:52
significant changes in human morphology
11:54
behavior and Technology occurred brain
11:58
volume increased sharp Lalo's tools
12:00
emerged and evidence of fire use and
12:03
complex settlements reflected enhanced
12:07
abilities this era was influenced by the
12:10
early to Middle Pline transition marked
12:13
by global climatic fluctuations
12:15
impacting ecosystems human populations
12:19
and their evolutionary
12:21
trajectory ancient DNA studies show
12:24
insights into relationships between
12:26
neanderthals denisovans and modern
12:29
humans modern humans diverge from
12:31
neanderthals around 500,000 years ago
12:34
and denisovans around 600,000 years
12:38
ago these findings illustrate diverse
12:41
evolutionary paths taken by human groups
12:44
over hundreds of thousands of years
12:47
highlighting the dynamic nature of human
12:51
worldwide between approximately 600,000
12:55
and 300,000 years ago different
12:57
populations of a ancestor began to show
13:01
variations leading to the formation of
13:05
groups these groups or subspecies were
13:08
geographically separated but still
13:10
belong to the same ancestral
13:13
species after 300,000 years ago new
13:17
species emerged from these
13:21
populations these new species were
13:23
defined based on genetic morphological
13:26
and behavioral differences known as
13:31
there's ongoing debate about how these
13:33
speciation events occurred in the
13:35
taxonomic status of many fossil
13:37
specimens found in Africa and
13:40
Eurasia the most curious question is the
13:43
exact identity of the last common
13:45
ancestor from which modern humans
13:47
neanderthals and Denny sens ultimately
13:51
originated most Scholars identify homo H
13:54
highle binus as the last common ancestor
13:57
based on fossil evidence
14:00
homo heidleberg insis existed between
14:02
approximately 900,000 and 300,000 years
14:05
ago and exhibited significant
14:07
variability in morphology and
14:10
geography in 1907 a Jawbone was found
14:14
near heidleberg Germany and the Sands
14:17
deposited by the necer river the species
14:19
was named homo Heidelberg insus popular
14:23
known as the Mau jaw it was well
14:25
preserved lacking a bony chin but with
14:28
teeth similar to those of modern
14:30
humans primitive features in the
14:33
specimen suggested that homo
14:34
heidelbergensis could be distinguished
14:36
from Asian and African representatives
14:40
erectus the Mau jaw is often grouped
14:44
with mandibles and partial craniums from
14:46
arago cave in France and petrona in
14:50
Greece if we include the SEMA deos Usos
14:53
in Spain additional skulls and numerous
14:56
postcranial bones can be added to the
15:00
typod the petrona and arago specimens
15:03
also share anatomical similarities with
15:06
crania from various locations including
15:09
broken Hill also called cab Cranium in
15:12
Zambia a lance Fontaine in South Africa
15:15
and Bodo from the middle aash of
15:18
Ethiopia this suggests that homo
15:21
heidleberg inses may have inhabited both
15:24
Europe the species is considered
15:27
potentially related not only to
15:29
neanderthals but also to modern
15:33
humans anatomically homo highle Beres
15:36
had a bigger brain compared to Homo
15:39
erectus averaging around
15:44
cm the skull relatively is taller than
15:48
erectus the front of the head is wider
15:51
the sides are high and arched like in
15:53
modern humans and the back is more
15:56
rounded the part at the back of the
15:58
skull is is longer than in Homo
16:01
erectus the jaw joint is similar to that
16:04
of homo sapiens along with certain parts
16:07
of the ear bone the Bony ridge above the
16:13
large the front part of the head is
16:15
flatter like in Homo
16:17
erectus the overall shape is less round
16:20
compared to modern humans the face is
16:23
connected to the skull in a way that
16:26
emphasizes the projection of the face
16:30
the nose opening is more vertical and
16:32
the canal in the front of the upper jaw
16:34
is placed more forward than in Homo
16:38
rectus the point where the two halves of
16:41
the jaw meat is receding and there's a
16:44
developed internal area for the teeth
16:47
European jaws of homo H highle bines
16:49
also look similar to
16:52
neanderthals some individuals from Sima
16:55
de los Usos were tall and robust with
16:58
limbo bones and a pelvis suggesting
17:00
possible differences in size between
17:02
males and females similar to what we see
17:07
populations the classification of homo
17:10
heidelbergensis in the Human family Tree
17:13
is a debated topic the Mau mandible
17:16
dating back 69,000 years is among the
17:19
oldest middle Pline fossils in Europe
17:22
aligning with findings in Bodo
17:25
Africa some scientists suggest replacing
17:28
homo heidelbergensis with homo boduan as
17:31
supporters say it clarifies middle ply
17:35
classification especially from Africa
17:39
Mediterranean opponents argue against
17:42
changing long-standing names expressing
17:45
concerns about confusion and potential
17:50
motivations the evolution and
17:52
classification of these middle plast
17:53
toine hominins is debated one view
17:56
groups populations into evolving grades
17:59
within Homo sapiens indicating a single
18:02
lineage with brain and skull
18:05
advancements alternatively some proposed
18:07
cladogenesis within homo arguing for a
18:10
unique morphology in living humans that
18:13
separates them from most fossil hominins
18:16
they believe only those sharing specific
18:18
features should be considered Homo
18:21
sapiens some researchers propose two
18:24
lineages the European Branch including
18:27
neanderthals traces back to the middle
18:29
plasto scene through fossils like
18:31
petralona arago seelos Usos and
18:36
Mau this perspective suggests repeated
18:39
speciation events another lineage rooted
18:42
in Africa includes specimens like broken
18:45
Hill elans fontine and Bodo referred to
18:50
rensis some late members display a mix
18:53
of archaic and modern
18:55
features an alternative view argues
18:58
minor morphological differences in
19:00
ancient European and African fossils
19:03
attributing them to geography and
19:06
variation this approach Advocates
19:08
classifying them under one taxon homo
19:11
hyal bensis challenging distinct
19:15
lineages if the broader homo
19:17
heidelbergensis classification is
19:19
accepted it could be the ancestor to
19:22
neanderthals and modern Homo sapiens
19:25
determining the preferred scenario
19:27
depends on the strength of morphometric
19:30
similarities notably findings from the
19:33
petrona cranium support homo H highle
19:36
bensis as a widespread polytypic
19:40
species around 1,000 to 900,000 years
19:44
ago earth's climate experienced glacial
19:47
Cycles every 100,000
19:49
years at the beginning of the middle pla
19:52
scene there was a notable change in the
19:55
environment marked by The Disappearance
19:57
of villafranchian faas leading to a
19:59
decline or disappearance of human
20:02
populations using oldo one
20:05
tools after about 650,000 years ago
20:09
there was a shift in Europe we start
20:11
seeing the use of more advanced Hulan
20:14
tools associated with human remains the
20:17
tools and fossils indicate similarities
20:21
contexts it suggests a possible
20:24
migration route from Africa to Europe
20:27
through the Middle East
20:29
in West Asia aulian Technologies
20:33
possibly originating in Africa appeared
20:36
before 1.35 million years ago in ubia
20:39
Israel and from 750,000 years ago
20:43
onwards in jeser Ben Yakov
20:47
Israel this aligns with archaeological
20:50
sites and human fossils in mid low
20:53
latitude regions of Africa and Europe
20:57
fossils from Mau and Germany and arago
20:59
cave in France distinct from homo
21:02
antecessor resemble those from terine in
21:05
Algeria suggesting homo heidleberg an's
21:08
spread from Africa to Eurasia
21:13
artifacts these findings including tools
21:16
and human remains in Italy and Frei
21:19
indicate advancement possibly influenced
21:21
by Africa through the West
21:23
Asia after reaching Europe in the middle
21:26
PTO scene homo heidleberg is adapted to
21:29
harsh conditions during glacial Cycles
21:32
leading to Neanderthal characteristics
21:34
through genetic drift in small or
21:38
groups concurrently homo heidelbergensis
21:40
in Africa evolve toward anatomically
21:43
modern Homo sapiens fossils from
21:46
locations like aoud in Morocco omo
21:49
kibish in Ethiopia heru in the middle
21:51
aash region and Loli in Tanzania
21:54
document this progression towards homo
21:58
scientists are debating recent findings
22:01
at s deos Usos challenging our
22:04
understanding of Neanderthal
22:05
Evolution the skulls from Sima delos
22:08
usus exhibit traits expected in early
22:12
neanderthals characteristics like a bony
22:15
ridge above the eyes broad nasal bones
22:18
and a distinct part at the back of the
22:19
head suggests similarities to
22:21
neanderthals teeth from SEMA deusas
22:24
closely resemble Neanderthal
22:26
teeth compared to other ancient skulls
22:29
like Mau or arago Calusa skulls appear
22:35
like some scientists propose Sima deos
22:38
Usos as the source population for
22:41
neanderthals while Mau and arago
22:43
represent a different
22:45
group this suggests two coexisting
22:48
species during the European Middle plene
22:51
homo heidelbergensis predating
22:52
neanderthals and early neanderthals
22:55
challenging previous Notions of human
23:00
fossils across Europe hint that
23:02
Neanderthal traits showed up first in
23:05
the Western and Atlantic regions and
23:08
later genetic signals also complicate
23:12
things mitochondrial DNA suggests a link
23:15
to denisovans while nuclear DNA shows
23:19
neanderthals this makes it hard to place
23:21
SEMA deos Usos with neanderthals
23:24
especially with denisovans in the
23:27
mix Homo sapiens likely originated in
23:30
Africa during the late middle
23:32
Pline potentially evolving from homo H
23:35
highle Bergens is due to climate
23:38
shifts following Homo erectus a gap in
23:42
fossil record spans 900 to 600,000 years
23:45
ago with Advanced features emerging
23:47
around 600,000 years ago mainly in
23:52
Africa further advancements including
23:55
larger brains and modern features occur
23:58
occurred around 300,000 years
24:01
ago some researchers thought homo
24:04
antecessor might be the common ancestor
24:06
of neandertals and modern humans because
24:08
it has a modern-like face however this
24:14
challenges the geographic position of
24:16
the Iberian Peninsula in the western
24:19
most extreme of the Eurasian continent
24:21
is far away from Africa where most
24:24
believe modern humans
24:26
originated moreover the geological age
24:29
of the td6 level is difficult to
24:32
conciliate with the fact that neander
24:34
paals and modern humans split later than
24:36
this age recent genetic studies suggest
24:40
the split between modern humans and
24:42
neandertals happened between
24:48
ago some researchers think modern like
24:51
facial features evolved independently in
24:54
different places and times homo
24:56
antecessor might not be the direct
24:58
ancestor of modern humans but rather a
25:01
side branch in Western
25:03
Europe however from a cladistic point of
25:06
view this lineage would stem out very
25:08
close to the last common ancestor of
25:10
homo sapiens and homon
25:14
neanderthalensis another population from
25:17
a nearby site called sadela lefante
25:20
dated to about 1.1 to 1.2 million years
25:23
ago might represent an earlier migration
25:26
into Europe though this is is still
25:29
uncertain studies of early human Dental
25:32
features suggest that no known fossil
25:34
including antecessor and heidleberg enes
25:37
fit the role last common ancestor of
25:40
neandertals and modern
25:42
humans there is a possibility that the
25:45
genetic estimates for the split between
25:47
neandertals and modern humans might be
25:50
too recent and differences and dental
25:52
features could have developed before
25:56
diverged homo intercessor does share
25:59
some traits with neandertals and other
26:01
early hominins but these traits might
26:04
not specifically indicate a direct
26:06
lineage instead they may reflect common
26:09
features in early human
26:12
populations ancient DNA analysis has
26:15
been tough because ancient DNA doesn't
26:19
time alternatively ancient proteins
26:22
found in Dental enamel are helping to
26:25
understand the position of antecessor
26:28
in a recent study scientists analyzed
26:31
Dental enamel from two key fossil groups
26:34
homo antecessor from Spain and Homo
26:38
Georgia extracting proteins from the
26:41
deines dentine sample only yielded a few
26:44
collagen fragments so they didn't
26:48
further however they successfully
26:50
retrieved and analyzed ancient enamel
26:53
proteins from both hominin and recent
26:57
teeth the the proteins included those
26:59
specific to enamel like amelogenin and
27:02
enamelin as well as some common proteins
27:05
like serum Alban and
27:07
collagen homo antecessor is closely
27:10
related to but not part of the group
27:13
that includes neanderthals denisovans
27:18
humans homo antecessor might be a sister
27:20
group to the common ancestor of these
27:23
species meaning it was closely related
27:25
but separate this aligns with other
27:28
findings that homo antecessor lived
27:30
around the time when Homo sapiens and
27:32
Neanderthals are thought to have
27:35
diverged it's thought that the ancestors
27:38
of neandertals and modern humans split
27:40
earlier than previously believed and
27:42
this split might have involved different
27:44
populations moving into
27:46
Europe there is evidence suggesting that
27:49
Eurasia played a significant role in
27:51
early human evolution with migrations
27:54
into Europe occurring due to various
27:56
Geographic and climatic
27:59
factors the current theory suggests that
28:02
modern humans originated in Africa while
28:05
neandertal evolved in
28:08
Europe however some believe that early
28:10
human populations might have mixed or
28:13
migrated between Africa and Eurasia
28:16
particularly through the letine corridor
28:18
key region connecting Africa and
28:22
Eurasia this mixing and migration could
28:24
have continued until climatic changes
28:27
around 1 million years ago which might
28:29
have contributed to the separation of
28:31
populations and the eventual evolution
28:34
of neandertals in modern humans