0:05
The origin of modern humans remains one
0:07
of the most complex and debated topics
0:10
in paleoanthropology.
0:12
During the middle to late pleaene
0:14
period, our ancestors shared the planet
0:17
with multiple other human species across
0:19
Africa and Eurasia, creating a
0:21
fascinating mosaic of human diversity.
0:25
These included the Neanderthalss in
0:27
Europe, the Denisovvens in Asia, the
0:30
smallrained but sophisticated Homonyi in
0:32
South Africa, the dimminionive hobbits
0:35
or homo floresensis in Indonesia, and
0:38
the recently discovered Homolusonensis
0:40
in the Philippines. This period
0:42
represents the last time Earth was home
0:44
to multiple human species before homo
0:47
sapiens became the sole survivor. The
0:50
relationships between these groups, who
0:52
evolved from whom, who interbred, and
0:54
who went extinct, continue to challenge
0:56
scientists as new discoveries emerge.
1:00
Asia, in particular, presents a crucial
1:02
yet puzzling piece of the story of human
1:05
origins. Fossils discovered across Asia,
1:08
including the Namada, Mara, Dali,
1:11
Ginusan, Shu Chang, Harbin, and Wongdong
1:14
specimens provide crucial insights into
1:17
human evolution during the middle to
1:19
late plea scene. However, poor
1:22
preservation of these fossils and
1:24
interpretations influenced by regional
1:26
continuity theories have made it
1:28
difficult to place them accurately in
1:30
human evolutionary history. The Namada
1:32
fossil discovered in the Namada Valley
1:35
of Madia Pradesh, India is considered
1:37
either a late homo erectus or an archaic
1:40
homo sapiens and is an important hominin
1:43
finds in South Asia. In China, several
1:46
key fossil sites have yielded remains
1:48
with a mosaic of archaic and modern
1:50
features. The Mara skull from Guangdong
1:53
province shows intermediate traits
1:54
between Homo erectus and homo sapiens.
1:57
The Darly cranium from Shan Xi and the
2:00
Ginyu Shan fossil from Liao Ning both
2:02
reflect transitional forms often
2:04
attributed to Homohidlebergensis or
2:06
early Homo sapiens dating to
2:08
approximately 200,000 to 260,000 years
2:11
ago. The Shu Chang fossils from Henan
2:15
province display a combination of traits
2:17
linked to both archaic humans and
2:19
possible Denisovven lineage. While the
2:22
Huong Dong fossils in Ani Province
2:25
dating to around 300,000 years ago show
2:28
a unique mix of primitive and modern
2:30
anatomical features. Together, these
2:32
discoveries highlight the complex
2:34
evolutionary landscape of early humans
2:36
across Asia and their importance for
2:39
understanding the emergence of Homo
2:41
sapiens. Among these, the Harbin cranium
2:44
stands out as the most complete and best
2:47
preserved because of its unique
2:49
combination of ancient and modern
2:51
traits. Scientists have proposed that it
2:53
belongs to a new human species named
2:56
Homongi. The species name Longi comes
2:59
from the Chinese term Long Jang, which
3:01
means dragon river, a poetic name for
3:03
the Hong Jang province, where the fossil
3:06
was found. The skull was discovered near
3:08
Dong Jang Bridge in Harbin City and was
3:11
later donated to the Geoscience Museum
3:13
at Hub Geo University in 2018.
3:16
The fossil was dated to about 146,000
3:20
years old, placing it in the late middle
3:22
pleaene period. What makes Homolonga
3:26
unique is its combination of features.
3:29
The skull is massive, larger than those
3:31
of most known ancient humans. It has a
3:34
large brain capacity 1420 ml similar to
3:39
modern humans and neanderals but the
3:42
shape of the brain case is long and low
3:44
not round like ours. The brow ridges are
3:48
thick and heavy. The face is wide but
3:51
flat and the eye sockets are large and
3:54
square. The nose is broad and the jaw
3:57
region suggests the presence of large
3:59
teeth but without a chin. When compared
4:03
to other fossil skulls from Asia and
4:05
Europe, Homo Longi shows clear
4:08
differences. It lacks the thick bone
4:10
structures of Homo erectus, the inflated
4:13
cheeks of Neanderthalss, and the sharply
4:15
angled back of the skull seen in
4:19
It also differs from the Darly and Huong
4:22
Dong skulls, which have their own
4:26
Because of this, scientists propose that
4:29
Darali and Hua Longdong belong to a
4:31
different species, Homo Daliansis, while
4:34
Harbin represents a completely new one,
4:39
Studies of ancient DNA from middle
4:41
pletoine sites in North Asia and Europe
4:43
have proven invaluable in establishing
4:45
the genetic relationships between early
4:47
humans and modern populations. One of
4:50
the key breakthroughs from this work was
4:52
the discovery of a previously unknown
4:55
archaic human population, the
4:58
identified through DNA extracted from
5:01
bones and teeth in Denisova cave in
5:03
Siberia. However, understanding
5:06
Denisovven physical characteristics
5:08
remains limited due to the
5:10
fragmentaryary nature of the fossils
5:12
which offer little morphological
5:14
information. Interestingly, the Harbin
5:17
skull appears to be closely related to a
5:19
jawbone found in Xiai, China, which is
5:22
believed to belong to the Denisovven
5:24
lineage. This connection raises the
5:27
possibility that the Harbin individual
5:29
may have also been part of the
5:30
Denisovven group or a close relative.
5:34
The Denisovvens were an ancient human
5:36
population that once roamed the Earth
5:39
alongside the Neanderals and modern Homo
5:41
sapiens. However, nobody knew about them
5:45
until a groundbreaking discovery in the
5:47
Denisan cave in Siberia.
5:50
In 2010, an analysis of mitochondrial
5:54
DNA extracted from a fingerbone found in
5:57
the Denise of a cave in the Russian Alai
5:59
revealed the existence of a new homminin
6:02
taxon. This newly described group was
6:04
found to be genetically distinct from
6:07
both Homo sapiens and homo
6:10
The mitochondrial DNA sequences showed
6:13
hapllet types outside the range of
6:15
variation of modern humans and
6:19
During the late middle to early upper
6:21
pletoine, three distinct groups emerged.
6:25
Early modern humans in Africa,
6:27
Neanderthalss in Europe, and Denisovvens
6:32
Molecular data suggests that the split
6:34
between the anderthals and Dennisovvens
6:36
occurred between 380 to 470,000 years
6:40
ago, while the branch leading to
6:42
Denisovvens and modern humans diverged
6:44
around 800,000 years ago.
6:49
migrated to Eurasia around 800,000 years
6:51
ago as indicated by the sites in Israel
6:54
in the Levant region.
6:57
Around 450,000 to 350,000 years ago,
7:01
Homohy Highlebergensis from the Levant
7:04
migrated to the Iranian plateau and
7:06
further to Central and North Asia
7:08
including Dennisova cave in southern
7:10
Siberia giving rise to Dennisovvens.
7:14
This migration marked the splitting of
7:16
late homohylebergensis into
7:18
Neanderthalss and denisovvens.
7:20
Moreover, late homohyleberlebergensis
7:23
from the Levant also reached to regions
7:25
such as Turkmenia, Kazakhstan and
7:28
Mongolia as indicated by the appearance
7:30
of Aulian industries in those areas.
7:33
Their likely route was north of the
7:34
Himalayas and Tibet. The mandible found
7:37
in Bishia cave on the northeastern
7:39
Tibetan plateau dates back to at least
7:42
160,000 years ago. The presence of
7:45
Dennisovvens in the region is also
7:47
supported by the adaptation of Tibetans
7:49
to high altitudes possibly through
7:52
introgression from denisovvens.
7:55
Hybridization events occurred between
7:57
these groups suggesting a meta
7:59
population that includes all three taxa.
8:02
The Alai region where Dennisova cave is
8:04
located was occupied by Neanderthalss,
8:07
Denisovvens and modern humans. Although
8:10
the precise timing and coexistence of
8:12
these groups remain uncertain,
8:15
the Denisova 11 girl represents an
8:17
example of first generation
8:19
hybridization between a Neanderthal
8:21
mother and a Denosovven father. Although
8:23
Dennisovvens likely lived across much of
8:25
Asia, scientists have only found direct
8:28
genetic evidence of them at a few
8:29
places. These include Dennisova cave in
8:32
Siberia, Bishia cave in Tibet, the
8:35
Penangu Channel near Taiwan, and
8:37
possibly a site in Laos called Tam Guuha
8:40
2 based on fossil shape. It's been
8:43
difficult to clearly classify
8:45
Dennisovvens because very few complete
8:47
bones or usable DNA samples have been
8:49
found. For example, a jawbone from
8:52
Bishia cave was linked to Dennisovvens
8:54
using protein analysis, but no DNA has
8:57
been recovered from it yet.
8:59
Interestingly, fossils from Sema de
9:02
loses in Spain have mitochondrial DNA
9:05
inherited from mothers similar to
9:07
Denisovvens, but their nuclear DNA
9:10
inherited from both parents is closer to
9:14
This suggests a complicated family tree
9:17
among ancient humans.
9:19
Most of the information about
9:21
Denisovvens has been inferred based on
9:23
analysis of modern human DNA. The
9:25
genetic diversity observed within the
9:27
Denisan lineage suggests their deep
9:29
divergence and separation into three
9:31
distinct branches across different
9:33
regions. The D0 lineage, often called
9:36
the Altidenisan lineage, is the best
9:39
studied and the only one directly
9:41
confirmed through ancient DNA
9:42
sequencing. It originates from Denisa
9:45
cave in the Alai Mountains of Siberia.
9:48
Multiple fossil remains have been
9:50
recovered from this cave and are
9:52
attributed to this group, including
9:54
Denisa 2, a juvenile mer genetically
9:57
sequenced and dated to over 100,000
9:59
years ago. Denisa 8, an adult mer dated
10:04
to around 120,000 years ago. Denisa 1920
10:09
and 21 show affinity to the older Ali
10:12
Denisven population and are dated to
10:14
roughly 160,000 to 200,000 years ago.
10:18
These individuals Denisa 2 8 19 2021
10:24
represent the earlier and more divergent
10:26
branch of Denisven in the Alai region.
10:29
By contrast, Denisa 3, a distal
10:32
fingerbone, and Denisa 4, another mer
10:36
younger, dated to about 50,000 to 76,000
10:39
years ago, and show greater genetic
10:41
similarity to Neanderls, possibly due to
10:44
increased ad mixture or regional
10:46
variation over time. These fossils
10:49
reflect the younger population of
10:50
Denisven in Denisa cave and suggest that
10:53
this site was occupied intermittently by
10:55
genetically distinct Denisven groups
10:57
over tens of thousands of years. Despite
11:00
the valuable DNA data, the D0 lineage
11:03
shows minimal genetic contribution to
11:05
modern human populations. This could
11:08
mean the Altidenisans had limited
11:10
contact with Homo sapiens or were
11:12
geographically isolated in Siberia.
11:15
Notably, Denisa 11 also belongs to this
11:18
lineage and reflects interactions
11:20
between archaic groups rather than
11:22
modern humans. The D1 Tibetan lineage is
11:25
genetically inferred from introgressed
11:27
Denisan DNA in modern East Asian
11:30
populations, particularly among Tibetans
11:35
This lineage is distinct from the Altai
11:37
Denisans and is thought to have lived in
11:39
mainland East Asia, especially in the
11:43
The strongest evidence for this lineage
11:45
is the EPAS 1 gene in Tibetans which
11:49
confers an advantage in high alitude low
11:51
oxygen environments. This gene's origin
11:54
cannot be traced to the altidenisans
11:56
confirming it stems from a different
11:58
diverged denisan population. The fossil
12:02
likely associated with this lineage is
12:04
the Bishia cast cave mandible from Shahi
12:06
Tibet dated to around 160,000 years ago.
12:17
The D2 lineage or oustrala papapuendis
12:21
also called oian andanan lineage is
12:24
based entirely on genetic introgression
12:26
patterns found in modern papuins
12:28
Aboriginal Australians and Melanesians
12:31
who carry up to 5% Denisven ancestry the
12:35
highest known levels globally
12:37
this lineage is deeply diverged from
12:39
both the alai and east Asian denisian
12:41
ans suggesting a long separated and
12:44
possibly isolated population of Denisans
12:47
living in island Southeast Asia.
12:49
Although no direct fossil evidence has
12:52
yet been tied to the D2 lineage, it is
12:54
possible that this group occupied parts
12:56
of Wallacea or the Sundealth Islands.
13:00
Speculative links have been made to
13:02
archaic fossils such as those at
13:04
Nagandong Java or even homo
13:07
flororosensis on Flores, but no genetic
13:10
data confirm a denis of anan identity.
13:12
This lineage is thought to have
13:14
interbred with homo sapiens migrating
13:16
through Southeast Asia towards Sahul,
13:19
prehistoric Australia New Guinea,
13:21
leaving a substantial genomic legacy in
13:27
The D3 lineage, Philippine Denisvens or
13:30
Negrito lineage, is genetically inferred
13:32
from indigenous Philippine populations,
13:35
especially the itamukon of northern
13:37
Luzon. These groups exhibit the highest
13:40
known proportion of Denisven DNA,
13:43
surpassing even Papwins. The D3 lineage
13:46
is deeply diverged from all other known
13:48
Denisven groups, implying a long
13:51
separate evolutionary trajectory likely
13:54
within the Philippine archipelago. The
13:56
populations of eastern Indonesians,
13:58
Papuans, Philippine Negritos, Siberians,
14:02
South Asians, and East Asians are among
14:04
the few existing groups that exhibit
14:06
substantial traces of genetic material
14:09
from Denisven. It is worth noting that
14:11
the region of islands Southeast Asia and
14:13
Papua, which consists of numerous
14:16
densely inhabited archipelos, holds some
14:19
of the earliest evidence of early
14:21
archaic humans outside Africa.
14:23
Additionally, it is believed that
14:25
archaic hominins like homo flororosensis
14:28
coexisted with modern humans in this
14:30
region. Modern Australian Aboriginals
14:33
and papwins have the highest proportion
14:35
of around 5 to 6% of Denisan ancestry
14:39
while American and mainland Asian
14:41
populations have a smaller proportion of
14:45
A possible fossil link to this lineage
14:49
discovered in Kalao cave Luzon and dated
14:53
to at least 67,000 years ago. Though DNA
14:56
could not be extracted, the fossils
14:58
primitive and unique features hint at
15:00
regional evolution of a Denisanlike
15:03
population. The high Denisan ancestry in
15:06
present-day Negrito groups supports the
15:09
idea that early homo sapiens arriving in
15:11
the Philippines interbred with an
15:13
indigenous Denisan population forming
15:16
the D3 lineages legacy. The harbin
15:19
cranium found in northeastern China has
15:22
been linked to the denisans through
15:23
ancient DNA analysis. Although
15:26
researchers couldn't extract DNA from
15:28
the bone or tooth, they successfully
15:30
retrieved it from dental calculus.
15:32
hardened plaque on the teeth. Out of 20
15:36
DNA samples prepared, seven contained
15:38
Denisven specific genetic markers and
15:41
none had Neanderal markers pointing to a
15:44
Denisven connection. To avoid bias,
15:47
scientists compared the harbin DNA not
15:50
just to modern humans, but to a broad
15:52
ancestral DNA reference built from
15:54
various ancient homins.
15:57
The resulting mitochondrial DNA sequence
16:00
closely resembled that of known
16:02
Denisvens, supporting the idea that the
16:04
Harbin individual belonged to or was
16:06
closely related to this mysterious human
16:11
Within the Denisven group, Harbin's
16:13
mitochondrial DNA is more similar to
16:15
older Denisvens, Denisa 2, 8, 19, 20,
16:19
and 21 than to the two younger ones,
16:22
Denise of a three and four, suggesting
16:25
an early divergence.
16:28
Earlier studies based on the skull shape
16:30
of Harbin had suggested it might belong
16:32
to a new species or be closely related
16:34
to Homo sapiens. But this new DNA study
16:38
shows that Harbin clearly falls within
16:40
the Denisven genetic range. Since the
16:43
Harbin skull is about 146,000 years old
16:46
and shares similar DNA, this suggests
16:49
Denisven lived widely across Asia during
16:51
the middle plea scene. This discovery is
16:54
important because it is the first time
16:56
Denisven DNA has been linked to physical
16:58
features in a wellpreserved fossil. It
17:01
may help scientists identify other
17:03
fossils like those from Dari, Jin Shan,
17:06
and Huong Dong as Denise of related even
17:10
if DNA isn't available by comparing bone
17:13
structure. However, the conclusion is
17:15
limited by the fact that only a small
17:17
amount of mitochondrial DNA was
17:19
recovered from harbin and no nuclear DNA
17:22
was found. The study of ancient DNA has
17:25
revealed significant genetic differences
17:27
between Denisvens and modern humans.
17:29
Denisven genomes differ from the
17:31
standard human genome by 11.7%
17:34
while the difference between
17:35
Neanderthalss and modern humans is
17:38
Denisvens and Neanderthalss are closely
17:41
related but distinct from homo sapiens.
17:44
Denisven show higher genetic diversity
17:47
than Neanderthalss but lower diversity
17:49
than modern humans indicating a larger
17:52
and more diverse population. Their
17:54
geographic distribution ranged from
17:56
North Asia to Southeast Asia. Denisven
18:00
carried genetic traits associated with
18:02
dark skin, brown hair, and brown eyes,
18:04
which are also present in modern humans.
18:07
Denisven likely played a role in
18:09
adaptive introgression, where early homo
18:12
sapiens acquired genes from Denisvens
18:14
that enhanced adaptation, disease
18:16
resistance, and immune systems. The
18:19
presence of human luccoite antigens in
18:21
modern humans inherited from Denise
18:24
supports this hypothesis.
18:27
For example, there is evidence that
18:29
adaptation to high altitude hypoxia
18:31
among modern-day Tibetans is a result of
18:34
introgression from Denisvens. There is
18:37
an argument that interbreeding between
18:39
humans and Denisovvens occurred east of
18:41
the Wallace's line, while another group
18:43
argues that interbreeding occurred
18:46
before humans crossed the Wallace's
19:04
The distribution of Denisven DNA in
19:06
modern populations suggests significant
19:09
interbreeding may have occurred after
19:11
early humans crossed Wallace's line.
19:14
Indigenous groups in New Guinea,
19:16
Australia, and nearby islands exhibit
19:19
the highest levels of Denisan ancestry,
19:24
Interestingly, populations in mainland
19:27
Asia, despite Denisven's historical
19:29
range, show lower levels of Denisan DNA.
19:33
This disparity indicates that Denise and
19:36
modern human interactions were
19:38
concentrated east of Wallace's line,
19:40
likely shaped by migration and
19:42
ecological boundaries.
19:45
Mainland populations may have
19:46
experienced a replacement effect where
19:49
later waves of East Asian populations
19:52
diluted or replaced Denisven DNA.
19:56
Even in isolated groups like the Andaman
19:58
Islanders who show no ad mixture with
20:00
other populations, no Denisven DNA has
20:03
been found. Denisven lived across both
20:07
mainland and island Southeast Asia,
20:10
thriving in diverse environments.
20:13
During the plea scene, when sea levels
20:16
dropped, vast savannah regions on the
20:18
sunda shelf, now submerged, likely
20:21
served as migration routes and refuges
20:23
for Denisvens during climate changes.
20:27
This mobility may have allowed them to
20:29
interact with modern humans in island
20:31
Southeast Asia, resulting in genetic
20:33
mixing. The only ancient human species
20:37
known to have crossed Wallace's line
20:39
before modern humans is homoresis.
20:43
This species remains mysterious as
20:46
scientists are unsure of its exact
20:48
evolutionary origins.
20:51
Some studies suggest it evolved from an
20:53
early ancestor of Homo erectus or
20:56
perhaps from an even older and more
20:58
primitive human species.
21:01
Evidence of stone tools on Flores dated
21:04
to over 1 million years ago indicates
21:06
that these homminins arrived on the
21:08
island very early. However, it's
21:11
difficult to link homo floresiensis to
21:15
The denisven had larger mers teeth which
21:19
don't match the anatomy of homop
21:22
Moreover, genetic studies suggest that
21:25
Denisven and modern humans shared a
21:27
common ancestor between 170,000 and 1
21:33
This timeline makes it unlikely that
21:35
homo floresensis which had been isolated
21:38
in the wisea region for so long was
21:41
directly related to the denisven.
21:44
The denisven on the other hand likely
21:46
arrived later around 600,000 years ago
21:50
and spread across a wider area of Asia
21:55
Some Denisven fossils may yet be
21:57
undiscovered, but possible candidates
21:59
could include the homolusencis from
22:02
Philippines or other mysterious human
22:04
fossils found in places like Narmmada
22:06
and Dari, Ginyu Shan, Mara, and Zujio,
22:12
These sites suggest that Denisven or
22:15
other ancient human species once had a
22:18
much broader range than previously
22:20
thought. Recently, evidence of human
22:23
activity has been found in the Kashmir
22:25
Valley, where around 300 to 400,000
22:28
years ago, humans relied on mammoth
22:31
carcasses, which further supports this
22:33
claim. The Dennisovven genome reveals
22:36
evidence of interbreeding with another
22:38
ancient population, but the identity of
22:41
this group remains a mystery. Scientists
22:44
speculate that Dennisovvens might have
22:46
mixed with older human species like Homo
22:49
erectus homo anticcessor or possibly a
22:52
late surviving homohyalbergis
22:55
in Asia. This raises intriguing
22:58
questions about how many ancient human
23:00
groups coexisted and interacted. The
23:04
fossil evidence from East Asia is
23:06
incomplete and scattered. But it hints
23:09
at a rich diversity of homminin species.
23:12
Discoveries like homo floresensis on
23:15
Flores and the homolusinesensis remains
23:17
from callo cave in the Luzon island of
23:20
Philippines suggest that multiple human
23:22
groups once inhabited wisaya.
23:25
This raises the possibility that some
23:28
early hominins might have even reached
23:30
New Guinea and Australia.
23:32
If true, it would mean ancient humans
23:35
were far more adaptable and capable of
23:38
long distance migration than previously
23:40
thought. Gene flow between Dennisovvens
23:44
and modern humans occurred primarily
23:46
east of Wallace's line, likely because
23:49
the circumstances of these encounters
23:51
were unique. Crossing Wallace's line
23:54
required watercraft, which suggests that
23:56
the first modern human groups reaching
23:58
Wallacea were small and isolated.
24:02
In such scenarios, interbreeding with
24:04
the established Dennisovven populations
24:06
may have been more likely as smaller
24:09
populations often rely on intergroup
24:11
mating to maintain genetic diversity.
24:15
Additionally, genetic signals from these
24:17
early interactions would be preserved
24:19
more easily in the descendants of such
24:21
small founding populations.
24:24
Interestingly, evidence suggests that
24:26
this gene flow was predominantly male
24:29
mediated, meaning Dennisovven males may
24:32
have interbred with modern human
24:34
females. This provides clues about the
24:37
nature of these interactions, though
24:39
much remains speculative.
24:42
Rapid dispersal into the tropical
24:44
regions of Wisa likely exposed early
24:47
modern humans to unfamiliar pathogens.
24:50
Hybridization with Dennisovvens who were
24:53
already adapted to the local environment
24:55
may have provided disease resistance
24:57
genes that offered a survival advantage.
25:01
This could have made interbreeding
25:02
beneficial and increased the likelihood
25:05
of these genetic traits being passed on
25:07
and preserved. As genomic research
25:10
progresses, it is expected to shed more
25:13
light on these interactions, revealing
25:15
how Neanderthal and Denisven DNA
25:18
contributed to the genetic makeup of
25:20
modern humans, and whether some of the
25:22
exchanged DNA had functional
25:24
significance, such as aiding in disease
25:28
resistance or environmental adaptation.
25:32
These discoveries are reshaping our
25:35
understanding of human evolution in
25:37
Asia, revealing it as a dynamic region
25:40
where multiple human species likely
25:45
The presence of Denisven across such a
25:47
wide geographic range from Siberia to
25:51
China to Southeast Asia suggests they
25:54
were a significant part of Asia's